The Neanderthals (English pronunciation /niˈændərˌθɔls/, /niˈændərˌtɔls/, /niˈændərˌtɑls/ or /neɪˈɑndərˌtɑls/) are a now-extinct species or subspecies within the genus Homo and closely related to modern humans. They are known from fossil specimens dating to the Pleistocene period and found in Europe and parts of western and central Asia. The term "Neanderthal", a shortening of "Neanderthal man", is sometimes spelled Neandertal, the modern spelling of the name of the Neander Valley in Germany where the species was first discovered.
Neanderthals are classified alternatively as a subspecies of Homo sapiens (Homo sapiens neanderthalensis) or as a separate species of the same genus (Homo neanderthalensis). The first proto-Neanderthal traits appeared in Europe as early as 600,000–350,000 years ago. Proto-Neanderthal traits are occasionally grouped with another phenetic species, such as Homo heidelbergensis, or a migrant form, Homo rhodesiensis.
The youngest Neanderthals include the Vindija Cave fossils in Croatia, which are between 33,000 and 32,000 years old. No definite specimens younger than 30,000 years ago have been found, but evidence of fire by Neanderthals in Gibraltar indicate they may have survived there until 24,000 years ago. Cro-Magnon or early modern human skeletal remains with "Neanderthal traits" were found in Lagar Velho (Portugal), dated to 24,500 years ago and interpreted as indications of extensively admixed populations.
Several cultural assemblages have been linked to the Neanderthals in Europe. The earliest, the Mousterian stone tool culture, dates to about 300,000 years ago. Late Mousterian artifacts were found in Gorham's Cave on the south-facing coast of Gibraltar. Other tool cultures associated with Neanderthal include Châtelperronian, Aurignacian, and Gravettian. These latter tool assemblages appear to have developed gradually within the populations, rather than being introduced by new population groups arriving in the region.
Neanderthal cranial capacity is thought to have been as large as that of modern humans, perhaps larger, indicating that their brain size may have been comparable, or larger, as well. In 2008, a group of scientists created a study using three-dimensional computer-assisted reconstructions of Neanderthal infants based on fossils found in Russia and Syria. The study showed Neanderthal and modern human brains were the same size at birth, but by adulthood, the Neanderthal brain was larger than the modern human brain. They were much stronger than modern humans, having particularly strong arms and hands. Males stood 164–168 cm (65–66 in) and females about 152–156 cm (60–61 in) tall.
Genetic evidence published in 2010 suggests they contributed DNA to anatomically modern humans, probably through interbreeding between Neanderthals and the earliest Humans that dispersed out of Africa. This is thought to have occurred between 80,000 and 50,000 years ago, shortly after (or perhaps before) the proto-Eurasians emigrated from Africa, while they were still one population. According to the study as much as 1–4% of the genome of the population that populated Eurasia was contributed by Neanderthals.
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